问题标题：
【StephenSpindlerStephenSpindlerandhiscolleaguesfromtheUniversityofCalifornia...的语法结够StephenSpindlerandhiscolleaguesfromtheUniversityofCaliforniaatRiversidehavefoundthatsomeofanelderlymouse'slivergenescanb】
问题描述：
StephenSpindlerStephenSpindlerandhiscolleaguesfromtheUniversityofCalifornia...的语法结够
StephenSpindlerandhiscolleaguesfromtheUniversityofCaliforniaat
Riversidehavefoundthatsomeofanelderlymouse'slivergenescanbemadeto
behaveastheydidwhenthemousewasyoungsimplybylimitingitsfoodforfour
weeks
canbemadeto怎么翻?
astheydid在这句话做什么结构
when后面是同位语么?
完整结构
马寿峰回答：
主语StephenSpindlerandhiscolleaguesfromtheUniversityofCaliforniaatRiverside
谓语havefound
宾语从句：thatsomeofanelderlymouse'slivergenescanbemadetobehaveastheydidwhenthemousewasyoungsimplybylimitingitsfoodforfourweeks
simplybylimitingitsfoodforfourweeks是补充说明的状语可以先不看
bemadetodosth表示被强迫（外力驱使下）去做某事
behave后面是从句的宾语部分如果要啰啰嗦嗦的写就是behavetodothethingsastheydowhentheyareyoung那些年纪大的老鼠会去做如他们年轻时做的事情（当然不是自发的,是被研究者驱使的
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